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A Simple Model for Leaf Placement

This Demonstration is a simple model of how leaves are placed on a plant stem. Sections of the cylindrical stem are unrolled into horizontal lines; the stem grows up the page. The red dots are where new leaves start to grow. The concentration of some hypothetical growth chemical is denoted by the curve at the top with varying shades of green. New leaves grow where the concentration of this chemical is the highest (i.e. lightest green) on the stem. This red dot is assumed to decrease the concentration around it, but overall concentration is increased before the next step. For more information, see Details.

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The angles between successive elements correspond to the horizontal distance between the two elements. Though these angles vary for the first few steps, there is, for most damping percentages, a rapid convergence to a fixed distance corresponding to an angle of almost exactly 137.5°.
The "damping percentage" refers to the damping of the memory of the previous position on the stem. When all memory is damped out (damping percentage = 100%), successive elements form 180° angles. Where there is very little damping, elements form at relatively random angles.
Phyllotaxis spirals are examples of structures whose successive elements are added at the golden ratio angle 137.5°.
For more information, see Stephen Wolfram's A New Kind of Science, pp. 400–412.
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