In age-structured models of populations, age classes or "cohorts" are introduced to distinguish individuals at different life stages. For

, where

is the number of different age classes, let

denote the number of individuals that are in cohort

at the discrete time (generation)

and assume an initial age distribution at

, here chosen as {0, 50%, 50%, 0, 0, 0}. In the population model put forward by P. H. Leslie ("On the Use of Matrices in Certain Population Mathematics,"
Biometrika,
33(3), 1945 pp. 183–212), the evolution from the given initial age distribution is determined by the recursion

for the distribution vector

and the "Leslie matrix"

. The parameters in

embody the survival rates yielding the fraction of the

class at time

that enter into the

class at time

, and the "fertility" or "fecundity" coefficients giving the average number of newborns to a member of the respective cohort. The age distributions displayed are computed as

, where

is the total population size at time

. It can be shown (by invoking the Perron–Frobenius theorem) that if

is regular then there exists an eigenvector with strictly positive components; this eigenvector yields the ''equilibrium" age distribution to which the evolution converges upon time

growing to infinity.